Thank you guys! i had not been able to find that paper. Lots of info there.
It does not seem to kill the animals nearly as quickly as I had thought. In fact, it seems that if there is enough food, they can reproduce even. Having seen limper herds, that came as a surprise.
I am also curious as to whether the limpers tend to congregate in certain areas (lowland, flat etc.)
Thank you for the information.